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Tomkins, J. L. and Hazel, W. N. in press. Explaining phenotypic diversity: The conditional strategy and threshold trait expression. In: Dung beetle Ecology and Evolution, Ed. Simmons, L. W. and Ridsdill-Smith, J. Wiley-Blackwell.
Tomkins, J. L., Penrose, M. A., Greeff, J. and LeBas N. R., 2010, Additive genetic breeding values correlate with the load of partially deleterious mutations Science: 328 892-894.
Tomkins, J. L., LeBas N. R., Witton, M. P., Martill, D. M. and Humphries, S. 2010, Positive allometry and the prehistory of sexual selection. The American Naturalist. 176: 141-148.
Tomkins, J. L. and Moczek, A.
P. 2009 Patterns
of threshold evolution in polyphenic insects under different developmental
models. Evolution 62:
459-468.
Kotiaho, J. S. LeBas, N. R., Puurtinen, M. and Tomkins, J. L. 2008 On the resolution of the lek paradox. Trends in Ecology and Evolution 23, 1-3.
Tomkins, J. L. & Hazel W. N. 2007 The status of the conditional evolutionarily stable strategy. Trends in Ecology and Evolution 22, 522-528.
Kotiaho, J. S.
LeBas, N. R., Puurtinen, M. and Tomkins, J. L. 2007 On female choice, heterozygosity and the lek
paradox. Animal Behaviour doi:10.1016/j.anbehav.2007.08.011
Simmons, L. W., Emlen, D. J. & Tomkins, J. L. 2007 Sperm competition games between sneaks and guards: a comparative analysis using dimorphic male beetles. Evolution 61, 2684–2692.
Tomkins, J. L., Kotiaho, J. S. & LeBas, N. R. 2006 Major differences in minor allometries. American Naturalist 167, 612-618.
Lukasik, P., Radwan. J., & Tomkins, J. L. 2006 Structural complexity of the environment affects the survival of alternative male reproductive tactics. Evolution 60, 399 – 403.
Gage,
M. J. G., Surridge, A. K., Tomkins, J. L., Green, E.,
Wiskin L.,
Tomkins, J. L., Kotiaho, J. S. & LeBas, N. R. 2005 Matters of scale: positive allometry and the evolution of male dimorphisms. American Naturalist. 165, 389-402.
Rutstein, A. N., Gilbert, L. & Tomkins, J. L. 2005 Experience counts: lessons from studies of differential allocation. Behavioral Ecology 16, 957-960.
Tomkins, J. L., Kotiaho, J. S. & LeBas, N. R. 2005 Phenotypic
plasticity in the developmental integration of morphological trade-offs and
secondary sexual trait compensation. Proceedings of the Royal Society of
Tomkins, J. L. & Brown, G. S. 2004 Population density drives the local evolution of a threshold dimorphism. Nature 431, 1099-1103.
Tomkins, J. L., Radwan, J., Kotiaho, J. S. & Tregenza, T. 2004 Genic capture and resolving the lek paradox. Trends in Ecology and Evolution 19, 323-328.
Tomkins, J. L., LeBas, N. R., Unrug, J. & Radwan, J. 2004. Testing the status-dependent ESS: population variation in fighter expression in the mite Sancassania berlesei. Journal of Evolutionary Biology 17, 1377-1388.
Knell, R. K., Pomfret, J. C. & Tomkins, J. L. 2004 The limits of elaboration: curved allometries reveal the
constraints on mandible size in stag beetles. Proceedings of the Royal
Society of
Unrug,
J., Tomkins, J. L. & Radwan, J. 2004 Alternative phenotypes and
sexual selection: can dichotomous handicaps honestly signal quality? Proceedings
of the Royal Society of
Tomkins, J. L. & Kotiaho, J. S. 2004 Publication bias in meta-analysis: seeing the wood for the trees. Oikos 104, 194-196.
Tomkins, J. L. & Simmons, L. W. 2003
Fluctuating asymmetry and sexual selection: paradigm shifts, publication bias
and observer expectation. In Developmental Instability: Causes and
Consequences (ed. M. Polak), pp. 231-261.
Kotiaho, J. S., Simmons, L. W., Hunt, J. & Tomkins, J. L. 2003 Males influence maternal effects that promote sexual selection. American Naturalist 161, 852-859.
Tomkins, J. L. & Simmons, L. W. 2002 Measuring relative investment: a case study of testes investment in species with alternative male reproductive tactics. Animal Behaviour 63, 1009-1016.
Kotiaho, J. S. & Tomkins, J. L. 2002 Meta-analysis can it ever fail ? Oikos 96, 551-553.
Radwan, J., Unrug, J. & Tomkins, J. L. 2002 Status dependence and morphological trade-offs in the expression of a sexually selected character in the mite, Sancassania berlesei. Journal of Evolutionary Biology 15, 744-752.
Colegrave, N., Kotiaho, J. S. & Tomkins, J. L. 2002 Mate choice or polyandry: reconciling genetic compatibility and good genes sexual selection. Evolutionary Ecology Research 2002, 911-917.
Kotiaho, J. S. & Tomkins, J. L. 2001 The discrimination of alternative male morphologies. Behavioral Ecology 12, 553-557.
Kotiaho, J. S., Simmons, L. W. & Tomkins, J. L. 2001 Towards a resolution of the lek paradox. Nature 410, 684-686.
Tomkins, J. L., Simmons, L. W. & Alcock, J. 2001 Brood provisioning strategies in Dawsons burrowing bee, Amegilla dawsoni (Hymenoptera: Anthophorini). Behavioral Ecology and Sociobiology 50, 81-89.
Tomkins,
J. L. & Kotiaho, J. S. 2001 Fluctuating asymmetry. In Encyclopedia of
Life Sciences.
Simmons, L. W., Tomkins, J. L. & Alcock, J. A. 2000 Can minor males of Dawson's burrowing bee, Amegilla dawsoni (Hymenoptera: Anthophorini) compensate for reduced access to virgin females through sperm competition ? Behavioral Ecology 11, 319-325.
Tomkins, J. L. & Simmons, L. W. 2000 Sperm competition games
played by dimorphic male beetles: fertilisation gains with equal mating
success. Proceedings of the Royal Society of
Tomkins, J. L. & Simmons, L. W. 1999 Heritability of size but not symmetry in a sexually selected trait chosen by female earwigs. Heredity 82, 151-157
Tomkins, J. L. 1999 The ontogeny of asymmetry in earwig forceps. Evolution 53, 157-163.
Simmons,
L. W., Tomkins, J. L., Kotiaho, J. S. & Hunt, J. 1999 Fluctuating
paradigm. Proceedings of the Royal Society of
Tomkins, J. L. 1999 Environmental and genetic determinants of the male forceps length dimorphism in the European earwig Forficula auricularia. L. Behavioral Ecology and Sociobiology 47, 1-8.
Simmons,
L. W., Tomkins, J. L. & Hunt, J. C. 1999 Sperm competition games
played by dimorphic male beetles. Proceedings of the Royal Society of
Kotiaho, J., S., Tomkins, J. L. & Simmons, L. W. 1999 Unfamiliar citations breed mistakes. Nature 400, 307.
Hunt, J., Kotiaho, J. S. & Tomkins, J. L. 1999 Dung pad residence time covaries with male morphology in the dung beetle Onthophagus taurus. Ecological Entomology 24, 174-180.
Tomkins, J. L., Simmons, L. W., Knell, R. J. & Norris, K. A. 1999 Correlates of ball size and rolling speed in the dung beetle Kheper nigroaeneus (Coleoptera : Scarabaeidae). Journal of Zoology 248, 483-487.
Tomkins, J. L. & Simmons, L. W. 1998 Female choice and manipulations of forceps size and symmetry in the European earwig Forficula auricularia L. Animal Behaviour 56, 347-356.
Tomkins, J. L. 1997 Sexual Selection
and the Evolution of Earwig Forceps: The
Simmons, L. W. & Tomkins, J. L. 1996 Sexual selection and the allometry of earwig forceps. Evolutionary Ecology 10, 97-104.
Tomkins, J. L. & Simmons, L. W. 1996 Dimorphisms and fluctuating asymmetry in the forceps of male earwigs. Journal of Evolutionary Biology. 9, 753-770.
Tomkins, J. L. & Simmons, L. W. 1995 Patterns of fluctuating
asymmetry in earwig forceps: no evidence for reliable signalling. Proceedings
of the Royal Society of
Simmons, L. W., Tomkins, J. L. & Manning, J. T. 1995 Sampling bias and fluctuating asymmetry. Animal Behaviour 49, 1697-1699.